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What's the practical purpose of changing calibers if it doesn't make a difference?
Changing calibers absolutely does make a difference. If it didn't, we wouldn't have so many. My comment about not shooting bullets harder has the implicit clarification that this is if it's chambered in the same caliber as another gun.
In their default factory configurations, the vast majority of AR-15's as well as the Mini 14 (the other gun pictured there) fire the same cartridge in the same caliber with approximately the same amount of energy, to no appreciable difference whatsoever from the point of view of whatever was shot with them. That is .223 Remington.
If you convert your gun to a different caliber, obviously the comparison no longer applies unless you compare it to other guns of the same caliber. But the Armalite platform is very modular, so making that conversion is super easy. This allows you to, just as an example, buy a bog standard model chambered in .223 and leave it that way for self defense or whatever, but then get an inexpensive .22LR upper to fire cheap .22LR ammo for target practice or plinking without having to spend the entire GDP of a third world country on ammunition, and/or keep a larger caliber receiver on hand in .300 Blackout or .450 Bushmaster or similar for hunting.
This saves you from having to buy and secure three separate guns for three separate tasks, especially considering you're unlikely to be needing all three at the same time. (I don't know about you, but I only have two hands.)
I think most gun owners tend to own quite a few guns. I also have seen where people tend to buy multiple AR-15 rifles in order to build something different every time for no discernable reason other than they like to build them and show them off. The issue is that the AR-15 platform attracts certain kinds of people who really don't have an interest in shooting as a sport. If it wasn't available I would guess that many of those people wouldn't buy some other rifle in its place.
I see you've never met the Ruger 10/22 grandpas. You want to talk about a bunch of guys who spend thousands of dollars buying, building, and ricing out rifles for "competition" or "varmint control" and inevitably have one or more builds they've never even fired nor do they ever intend to.
But it's got a rainbow-stained burl walnut thumbhole stock, magazine release lever conversion, 2" thick carbon fiber bull barrel, all stainless hardware, a $900 trigger group, 50 round aftermarket banana mag, a bipod, and a 10-32x240mm illuminated reticle night vision scope! You don't understand, I had to spend $8000 on building it because .22 ammo is just so cheap!
Some weirdos are just like that.